Round 4: Tossup 5

This parameter is plotted along a line running from mu to A in a “geometric” method developed by Alan Grafen. This parameter and competition are assumed to be locally enhanced for a viscous population in a method of estimating this parameter named for Goodnight and Queller. According to one theory, unusual asymmetries in this parameter within a population drive the evolution of eusociality. A less-than-average “negative” form of this parameter often arises from the emergence of (*) spiteful genes. One behavior is favored (-5[1])if (-5[1])“this parameter times benefit minus cost (10[1])is greater than zero,” per Hamilton’s rule. In a pedigree, this parameter equals one-eighth for cousins and one-half (10[1])for full-siblings. For 10 points, what parameter denoted r measures the genetic similarity between individuals? ■END■ (10[2])

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